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On the Origin of Species · ¶311

We should be extremely cautious in concluding that an organ could not have been formed by transitional gradations of some kind. Numerous cases could be given amongst the lower animals of the same organ performing at the same time wholly distinct functions; thus the alimentary canal respires, digests, and excretes in the larva of the dragon-fly and in the fish Cobites. In the Hydra, the animal may be turned inside out, and the exterior surface will then digest and the stomach respire. In such cases natural selection might easily specialise, if any advantage were thus gained, a part or organ, which had performed two functions, for one function alone, and thus wholly change its nature by insensible steps. Two distinct organs sometimes perform simultaneously the same function in the same individual; to give one instance, there are fish with gills or branchiæ that breathe the air dissolved…
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Charles Darwin · John Murray, London, 1859, first edition · Gutenberg #1228; original diagram from Darwin Online F373
Charles Darwin · John Murray, London, 1859, first edition · Gutenberg #1228; original diagram from Darwin Online F373
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