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On the Origin of Species · ¶414

The foregoing rules and facts, on the other hand, appear to me clearly to indicate that the sterility both of first crosses and of hybrids is simply incidental or dependent on unknown differences, chiefly in the reproductive systems, of the species which are crossed. The differences being of so peculiar and limited a nature, that, in reciprocal crosses between two species the male sexual element of the one will often freely act on the female sexual element of the other, but not in a reversed direction. It will be advisable to explain a little more fully by an example what I mean by sterility being incidental on other differences, and not a specially endowed quality. As the capacity of one plant to be grafted or budded on another is so entirely unimportant for its welfare in a state of nature, I presume that no one will suppose that this capacity is a specially endowed quality, but will…
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Charles Darwin · John Murray, London, 1859, first edition · Gutenberg #1228; original diagram from Darwin Online F373
Charles Darwin · John Murray, London, 1859, first edition · Gutenberg #1228; original diagram from Darwin Online F373
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